What is the meaning of ?

Formed the third-person singular indicative past definite, for back-vowel verbs. For the full paradigm, see the usage template.

Combining form of -a (possessive suffix) before most case suffixes, except that of the essive-formal.

of (used with a noun)

Plural suffix (with a-declension nouns)

Causative (valence-increasing) suffix

Oblique case suffix (with a-declension nouns)

forms the third-person singular future indicative form of verbs

Apocopic form of -ar; used preceding the pronouns lo, la, los or las

Alternative form of -ar in any position

Source: wiktionary.org